I like the idea of life and death being a central motivation of the conflict between these two. A classic battle between two forces of life are birth and death. Which brings to mind an image of a baby fighting a skeleton. Nahh, lets not go there (but maybe come back to that notion later...).
Life is a cycle, with birth, death and rebirth. Perhaps this battle can take place between agents of Birth and Rebirth respectively. Here we can introduce the notion of belief; one opponent wants to preserve life in its current form and the other wishes to hasten life's rebirth into a newer form, through death. And the force which brings them both together with the common purpose of protecting life utterly? A force of destruction, in its purest form seems efficient.
So, we have two beings, fighting in the name of different beliefs/philosophies but with ultimately the same purpose, coming together to combat something which threatens both.
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